Started over, same everything, except this time the AIs weren't random, but the exact same as last time.
Two things of note: My scouts have a weird auto-explore fixation on Bartledan, a planet two hops to my west. Noticed this with mk 1 scouts always following the route, decided to see what happened with a Mark IV scout starship or two. They park right on that wormhole for some reason. I took the other one manually hunting for the AI Homeworlds so I had something to look at.
Fortresses and AI Command Stations in the two adjacent systems without co-processors are covered with Shield Bearers, and the Mark 3 Fortresses are parked directly overhead. This influenced my decision not to pop any command stations at all. At the base 20 AIP and 31 minutes, the three adjacent worlds are at 522, 425, 488
If you'd like, I can compare with two vanilla AIs.
Correction: I have spire civilian leaders on, that means I'm actually 10 above base AIP.